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Please note that the content of this book primarily consists of articles available from Wikipedia or other free sources online. Theodor Boveri originally observed the fact that abnormal nuclear morphologies commonly occurred in cancer. Micronuclei are also referred to Howell-Jolly bodies; founded by hematologists William Henry Howell and Justin Marie Jolly in erythrocytes. Micronuclei induction by a chemical was first reported in Ehrlich ascites tumor cells treated with colchicine. The affect of environmental stressors on the expression of micronuclei was first analyzed in root tip cells under ionizing radiation.