High Quality Content by WIKIPEDIA articles! Professor George Christopher Williams (b. May 12, 1926) is an American evolutionary biologist. Williams is a professor emeritus of biology at the State University of New York at Stony Brook. He is best known for his vigorous critique of group selection. The work of Williams in this area, along with W. D. Hamilton, John Maynard Smith and others led to the development of a gene-centric view of evolution in the 1960s. Williams' 1957 paper Pleiotropy, Natural Selection, and the Evolution of Senescence is one of the most influential in 20th century evolutionary biology, and contains at least 3 foundational ideas. The central hypothesis of antagonistic pleiotropy remains the prevailing evolutionary explanation of senescence. In this paper Williams was also the first to propose that senescence should be generally synchronized by natural selection. According to this original formulation