Mary Leakey was a British archaeologist and anthropologist, who discovered the first skull of a fossil ape on Rusinga Island and also a noted robust Australopithecine called Zinjanthropus at Olduvai. For much of her career she worked together with her husband, Louis Leakey, in Olduvai Gorge, uncovering the tools and fossils of ancient hominines. She developed a system for classifying the stone tools found at Olduvai. She also discovered the Laetoli footprints. In 1960 she became director of excavation at Olduvai and subsequently took it over, building her own staff. After the death of her husband she became a leading palaeoanthropologist, helping to establish the Leakey tradition by training her son, Richard, in the field.