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High Quality Content by WIKIPEDIA articles! In spatial ecology and macroecology, scaling pattern of occupancy (SPO), also known as the area-of-occupancy is the way in which species distribution changes across spatial scales. In physical geography and image analysis, it is similar to the modifiable areal unit problem. Simon A. Levin (1992), states that the problem of relating phenomena across scales is the central problem in biology and in all of science. Understanding the SPO is thus one central theme in ecology.