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Please note that the content of this book primarily consists of articles available from Wikipedia or other free sources online. The Genetical Evolution of Social Behaviour is a 1964 scientific paper by the British evolutionary biologist W.D. Hamilton in which he mathematically lays out the basis for inclusive fitness. Hamilton, then only a PhD student, completed his work in London. It was based on Haldane''s idea, but Hamilton showed that it applied to all gene frequencies. Although initially obscure, it is now highly cited in biology books, and has gone on to reach such common currency that citations are now often unnecessary as it is assumed that the reader is so familiar with kin selection and inclusive fitness that he need not use the reference to obtain further information.