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In this study, nine florules from the Chloropagus Formation near Fernley, Nevada, are dated at 14.7-13.4 million years. The author finds that dominant mixed conifer forest and sclerophyll woodland species of the Sierra Nevada-Klamath region replaced exotic deciduous hardwoods in the two lowest sites. He concludes that this change reflects the loss of adequate summer rain as upwelling from a colder ocean resulted from spreading East Antarctic ice.