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Please note that the content of this book primarily consists of articles available from Wikipedia or other free sources online. In enzymology, an UDP-glucose 4-epimerase (EC 5.1.3.2) is an enzyme that catalyzes the chemical reaction. UDP-glucose = UDP-galactose. Hence, this enzyme has one substrate, UDP-glucose, and one product, UDP-galactose. This enzyme belongs to the family of isomerases, specifically those racemases and epimerases acting on carbohydrates and derivatives. This enzyme participates in galactose metabolism and nucleotide sugars metabolism. It employs one cofactor, NAD+.